Professional Biography

I teach courses in Genetics and Evolutionary Biology, Neurobiology, and Immunology. I believe that students learn best by doing, so my courses emphasize problem-solving, data analysis, and student-designed lab projects.

In my research students and I study development of the vertebrate nervous system, with a particular focus on intercellular communication between developing nerve cells and their target tissues.

Academic Credentials

Ph.D. 1975, M.Phil. 1974 Columbia University

B.A. 1967 Johns Hopkins University